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The silver dapple gene, also known as the "Z" gene, is a gene that dilutes only the black on a horse. It will typically dilute a black mane and tail to flaxen, and a black body to a shade of brown or chocolate. Red based horses (chestnut, palomino, cremello) may carry the gene, but will not express it. They can therefore pass it on to their offspring, but the silver dapple gene just won't be visible in their own body color.
http://en.wikipedia.org/wiki/Silver_dapple_gene
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